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Anatomy

  • This is the second cranial nerve (CN II) and is the nerve of sight.

Skeletal Muscle:  1-40 cm long fibres, 10- 60 µm thick, according to myoglobin content there are:

Red fibres: lots of myoglobin, many mitochondriam slow twitching - tire slowly

White fibres:  less myoglobin,  less mitochondria, fast twitching - tire quickly

Intermediate fibres:

mixture of 2 above

Most muscles have all three - in varying ratios

Structure of skeletal muscle:

Light Microscopy:  Many nuclei - 35/mm,  Nuclei are oval - situated peripheral,  Dark and light bands

Electron Microscopy: Two types of myofilaments

Actin

- 5,6 nm

 3 components:

 -actin monomers,  

 -tropomyosin - 7 actin molecules long

- troponin

 actin monomers form 2 threats that spiral

- tropomyosin - lie in the groove of the spiral

- troponin - attach every 40 nm

- one end attach to the Z line

- other end goes to the middle of the sarcomere

- Z line consists of á actinin

Myosin:

- 15 nm

- 1,6 µm long

- The molecule has a head and a tail

- tails are parallel

- heads project in a spiral

- in the middle is a thickening

- thin threats bind the myosin at thickening (M line)

Contraction:

A - band stays the same, I - band, H - bands become narrower

Myosin heads ratchet on the actin molecule

Sarcolemma: 9 nm thick,  invaginate to form T-tubule,

 myofibrils - attach to the sarcolemma

Sarcoplasmic Reticulum:

specialized smooth EPR,  Consists of T-tubules, terminal sisternae and sarcotubules

It is speculated that there are gap junctions between the T-tubule and terminal sisterna

An impulse is carried into the fiber by the T-tubule from where it goes to the rest of the sarcoplasmic reticulum

Connective tissue coverings of the muscle

Endomycium around fibres, perimycium around bundles and epimycium around the whole muscle

Blood vessels and nerves in CT

CT goes over into tendon or aponeurosis which attaches to the periosteum

Nerves:

The axon of a motor neuron branches and ends in motor end plates on the fiber

Specialized striated fibres called spindles (stretch receptors) form sensory receptors in muscles telling the brain how far the muscle has stretched

Initially, four clefts exist; however, only one gives rise to a definite structure in adults.

1st pharyngeal cleft

Penetrates underlying mesenchyme and forms EAM.  The bottom of EAM forms lateral aspect of tympanic cavity.

2nd pharyngeal cleft

Undergoes active proliferation and overlaps remaining clefts.  It merges with ectoderm of lower neck such that the remaining clefts lose contact with outside.  Temporarily, the clefts form an ectodermally lined cavity, the cervical sinus, but this disappears during development.

MUSCLE

Types:

Skeletal (voluntary)
Cardiac (involuntary)
Smooth (involuntary)

Pharyngeal Arch

Arch Artery

Cranial Nerve

Skeletal elements

Muscles

1

Terminal Branch of maxillary artery

Maxillary and mandibular division of trigemenial (V)

Derived from arch cartilages (originating from neural crest):

From maxillary cartilages:

Alispenoid, incus

From mandibular:

Mackel’s cartilage, malleus

 

Upper portion of external ear (auricle) is derived from dorsal aspect of 1st pharyngeal arch.

 

Derived by direct ossification from arch dermal mesenchyme:

Maxilla, zygomatic, squamous portion of temporal bone, mandible

 

Muscles of mastication (temporalis, masseter, and pterygoids), mylohyoid, anterior belly of digastric, tensor tympani, tensor veli palatini (originate from cranial somitomere 4)

2

Stapedius artery (embryologic) and cortiotympanic artery (adult)

Facial nerve (VII)

Stapes, styloid process, stylohyoid ligament, lesser horns and upper rim of hyoid (derived from the second arch cartilage; originate from neural crest).

 

Lower portion of external ear (auricle) is derived from 2nd pharyngeal arch.

Muscles of facial expression (orbicularis oculi, orbicularis oris, auricularis, platysma, fronto-ooccipitalis, buccinator), posterior belly of digastric, stylohyoid, stapedius (originate from cranial somitomere 6)

3

Common carotid artery, most of internal carotid

Glossopharyngeal (IX)

Lower rim and greater horn of hyoid (derived from the third arch cartilage; originate from neural crest cells)

Sytlopharyngeus (originate from cranial somitomere 7)

4

Left: Arch of aorta;

Right: Right subclavian artery;

Original sprouts of pulmonary arteries

Superior laryngeal branch of vagus (X)

Laryngeal cartilages (Derived from the 4th arch cartilage, originate from lateral plate mesoderm)

Constrictors of pharynx, cricothyroid, levator veli palatine (originate from occipital somites 2-4)

6

Ductus arteriosus; roots of definitive pulmonary arteries

Recurrent laryngeal branch of vagus (X)

Laryngeal cartilages (derived from the 6th-arch cartilage; originate from lateral plate mesoderm)

Intrinsic muscles of larynx (originate from occipital somites 1 and 2)

The Soft Palate

  • This is the posterior curtain-like part, and has no bony support. It does, however, contain a membranous aponeurosis.
  • The soft palate, or velum palatinum (L. velum, veil), is a movable, fibromuscular fold that is attached to the posterior edge of the hard palate.
  • It extends posteroinferiorly to a curved free margin from which hangs a conical process, the uvula (L. uva, grape).
  • The soft palate separates the nasopharynx superiorly and the oropharynx inferiorly.
  • During swallowing the soft palate moves posteriorly against the wall of the pharynx, preventing the regurgitation of food into the nasal cavity.
  • Laterally, the soft palate is continuous with the wall of the pharynx and is joined to the tongue and pharynx by the palatoglossal and palatopharyngeal folds.
  • The soft palate is strengthened by the palatine aponeurosis, formed by the expanded tendon of the tensor veli palatini muscle.
  • This aponeurosis attaches to the posterior margin of the hard palate.

 
Anterior 2/3 of tongue Posterior 1/3 of tongue
Motor Innervation All muscles by hypoglossal nerve (CN XII) except palatoglossus muscle (by the pharyngeal plexus)
General Sensory Innervation
Lingual nerve (branch of mandibular nerve CN V3) Glossopharyngeal nerve (CN IX)
Special Sensory Innervation
Chorda tympani nerve (branch of facial nerve) Glossopharyngeal nerve (CN IX)

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