The mixed dentition

I. Transition dentition between 6 and 12 years of age with primary tooth exfoliation and permanent tooth eruption

2. Its characteristic features have led this to be called the ugly duckling stage because of

a. Edentulated areas

b. Disproportionately sized teeth

c. Various clinical crown heights

d. Crowding

e. Enlarged and edematous gingiva

f. Different tooth colors

Enamel

Structural characteristics and microscopic features

a.  Enamel rods or prisms

(1) Basic structural unit of enamel.

(2) Consists of tightly packed hydroxyapatite crystals. Hydroxyapatite crystals in enamel are four times larger and more tightly packed than hydroxyapatite found in other calcified

tissues (i.e., it is harder than bone).

(3) Each rod extends the entire thickness of enamel and is perpendicular to the dentinoenamel junction (DEJ).
 

b. Aprismatic enamel

(1) The thin outer layer of enamel found on the surface of newly erupted teeth.

(2) Consists of enamel crystals that are aligned perpendicular to the surface.

(3) It is aprismatic (i.e., prismless) and is more mineralized than the enamel beneath it.

(4) It results from the absence of Tomes processes on the ameloblasts during the final stages of enamel deposition.

c. Lines of Retzius (enamel striae)

(1) Microscopic features

 (a) In longitudinal sections, they are observed as brown lines that extend from the DEJ to the

tooth surface.

 (b) In transverse sections, they appear as dark, concentric rings similar to growth rings in a tree.
 

(2) The lines appear weekly during the formation of enamel.
 

(3) Although the cause of striae formation is unknown, the lines may represent appositional or incremental growth of enamel. They may also result from metabolic disturbances of ameloblasts.

(4) Neonatal line

(a) An accentuated, dark line of Retzius that results from the effect of physiological changes

on ameloblasts at birth.

(b) Found in all primary teeth and some cusps of permanent first molars.

d. Perikymata

(1) Lines of Retzius terminate on the tooth surface in shallow grooves known a perikymata.

(2) These grooves are usually lost through wear but may be observed on the surfaces of developing teeth or nonmasticatory surfaces of formed teeth.
 

e. Hunter-Schreger bands

(1) Enamel rods run in different directions. In longitudinal sections, these changes in direction result in a banding pattern known as HunterSchreger bands.

(2) These bands represent an optical phenomenon of enamel and consist of a series of  alternating dark and light lines when the section is viewed with reflected or polarized

light.

f. Enamel tufts

(1) Consist of hypomineralized groups of enamel rods.

(2) They are observed as short, dark projections found near or at the DEJ.

(3) They have no known clinical significance.

g. Enamel lamellae
 

(1) Small, sheet-like cracks found on the surface of enamel that extend its entire thickness.

(2) Consist of hypocalcified enamel.

(3) The open crack may be filled with organic material from leftover enamel organ components, connective tissues of the developing tooth, or debris from the oral cavity.

(4) Both enamel tufts and lamellae may be likened to geological faults in mature enamel.
 

h. Enamel spindle
 

(1) Remnants of odontoblastic processes that become trapped after crossing the DEJ during the differentiation of ameloblasts.
 

(2) Spindles are more pronounced beneath the cusps or incisal edges of teeth (i.e., areas where occlusal stresses are the greatest).
 

FORMATION OF THE PERMANENT DENTITION

Twenty deciduous tooth buds are formed initially.
Proliferative activity of the dental lamina during the bell stage that leads to formation of permanent tooth buds (cap stage) lingual of each deciduous tooth germ.
Molars have no predecessors; they are formed by posterior proliferation of the dental lamina.

HARD TISSUE FORMATION

Hard tissue formation starts at the late stages of the bell stage.
Differentiatioin of cells into odontoblasts and ameloblasts.
The cells of the inner dental epithelium will become ameloblasts.
The cells of the dental papilla opposite to the inner dental epithelium will become odontoblasts.
Dentin is formed before enamel.
Dentin initiates the formation of enamel.

ROOT FORMATION

The root of the tooth is composed by dentin and cementum.
Dentinogenesis is initiated by the odontoblasts.
Odontoblasts are formed as epithelial cells continue to proliferate from the cervical loop as a double layer of cells known as Hertwig's root sheath.

TOOTH SHAPE

The shape of the crowns results from the interaction of inner dental epithelium and the dental papilla.
The cells of the inner dental epithelium have a programmed proliferation.
This internal program determines the tooth form.

The fate of the dental lamina

Rests of Serres
The rest of Serres are rests of the dental lamina identified in the gingival soft tissues.
They are round to ovoid aggregates of epithelial cells that have clear cytoplasm (glucogen rich).
They result from early breakup of the dental lamina during bell stage.

Rests of Malassez
The rests of Malassez result from breakup of the Hertwig's root sheath during root formation.
They can be identified in the periodontal ligament and are responsible for the development of radicular cysts.

Age changes in the dentition

I. After the teeth have reached full occlusion, microscopic tooth movements occur to compensate for wear at the contact area (Mesial Drift) and occlusal surfaces (by Deposition of cementum at the root apex)

2. Attrition of incisal ridges and cusp tips may be so severe that dentin may become exposed and intrinsically stained

3. Secondary dentin may be formed in response to dental caries, trauma, and aging and result in decreased pulp size and tooth sensation

MANDIBULAR SECOND MOLAR

Facial: When compared to the first molar, the second molar crown is shorter both mesiodistally and from the cervix to the occlusal surface. The two well-developed buccal cusps form the occlusal outline. There is no distal cusp as on the first molar. A buccal developmental groove appears between the buccal cusps and passes midway down the buccal surface toward the cervix.

Lingual: The crown is shorter than that of the first molar. The occlusal outline is formed by the mesiolingual and distolingal cusps.

Proximal: The mesial profile resembles that of the first molar. The distal profile is formed by the distobuccal cusp, distal marginal ridge, and the distolingual cusp. Unlike the first molar, there is no distal fifth cusp.

Occlusal: There are four well developed cusps with developmental grooves that meet at a right angle to form the distinctive "+" pattern characteristic of this tooth.

Contact Points; When moving distally from first to third molar, the proximal surfaces become progressively more rounded. The net effect is to displace the contact area cervically and away from the crest of the marginal ridges.

Roots:-The mandibular second molar has two roots that are smaller than those of the first molar. When compared to first molar roots, those of the second tend to be more parallel and to have a more distal inclination.

MANDIBULAR SECOND BICUSPID

Facial: From this aspect, the tooth somewhat resembles the first, but the buccal cusp is less pronounced. The tooth is larger than the first.

Lingual: Two significant variations are seen in this view. The most common is the three-cusp form which has two lingual cusps. The mesial of those is the larger of the two. The other form is the two-cusp for with a single lingual cusp. In that variant, the lingual cusp tip is shifted to the mesial.

Proximal: The buccal cusp is shorter than the first. The lingual cusp (or cusps) are much better developed than the first and give the lingual a full, well-developed profile.

Occlusal: The two or three cusp versions become clearly evident. In the three-cusp version, the developmental grooves present a distinctive 'Y' shape and have a central pit. In the two cusp version, a single developmental groove crosses the transverse ridge from mesial to distal

Contact Points; Height of Curvature: From the facial, the mesial contact is more occlusal than the distal contact.The distal marginal ridge is lower than the mesial marginal ridge

Root Surface:-The root of the tooth is single, that is usually larger than that of the first premolar  

the lower second premolar is larger than the first, while the upper first premolar is just slightly larger than the upper second

There may be one or two lingual cusps