Crown stage

Hard tissues, including enamel and dentin, develop during the next stage of tooth development. This stage is called the crown, or maturation, stage by some researchers. Important cellular changes occur at this time. In prior stages, all of the inner enamel epithelium cells were dividing to increase the overall size of the tooth bud, but rapid dividing, called mitosis, stops during the crown stage at the location where the cusps of the teeth form. The first mineralized hard tissues form at this location. At the same time, the inner enamel epithelial cells change in shape from cuboidal to columnar. The nuclei of these cells move closer to the stratum intermedium and away from the dental papilla.

The adjacent layer of cells in the dental papilla suddenly increases in size and differentiates into odontoblasts, which are the cells that form dentin. Researchers believe that the odontoblasts would not form if it were not for the changes occurring in the inner enamel epithelium. As the changes to the inner enamel epithelium and the formation of odontoblasts continue from the tips of the cusps, the odontoblasts secrete a substance, an organic matrix, into their immediate surrounding. The organic matrix contains the material needed for dentin formation. As odontoblasts deposit organic matrix, they migrate toward the center of the dental papilla. Thus, unlike enamel, dentin starts forming in the surface closest to the outside of the tooth and proceeds inward. Cytoplasmic extensions are left behind as the odontoblasts move inward. The unique, tubular microscopic appearance of dentin is a result of the formation of dentin around these extensions.

After dentin formation begins, the cells of the inner enamel epithelium secrete an organic matrix against the dentin. This matrix immediately mineralizes and becomes the tooth's enamel. Outside the dentin are ameloblasts, which are cells that continue the process of enamel formation; therefore, enamel formation moves outwards, adding new material to the outer surface of the developing tooth.

Clinical importance of cementum

1) Deposition of cementum continues throughout life.
The effects of the continuous deposition of cementum are the maintenance of total length of the tooth (good) and constriction of the apical foramen (bad).
2) With age, the smooth surface of cementum becomes more irregular due to calcification of some ligament fiber bundles. This is referred to as spikes.

Behavior of cementum in pathologic conditions

Cap stage

The first signs of an arrangement of cells in the tooth bud occur in the cap stage. A small group of ectomesenchymal cells stops producing extracellular substances, which results in an aggregation of these cells called the dental papilla. At this point, the tooth bud grows around the ectomesenchymal aggregation, taking on the appearance of a cap, and becomes the enamel (or dental) organ. A condensation of ectomesenchymal cells called the dental follicle surrounds the enamel organ and limits the dental papilla. Eventually, the enamel organ will produce enamel, the dental papilla will produce dentin and pulp, and the dental follicle will produce all the supporting structures of a tooth

MAXILLARY SECOND MOLAR

The second molars are often called 12-year molars because they erupt when a child is about 12 years

Facial: The crown is shorter occluso-cervically and narrower mesiodistally whe compared to the first molar. The distobuccal cusp is visibly smaller than the mesiobuccal cusp. The two buccal roots are more nearly parallel. The roots are more parallel; the apex of the mesial root is on line with the with the buccal developmental groove. Mesial and distal roots tend to be about the same length.

Lingual: The distolingual cusp is smaller than the mesiolingual cusp. The Carabelli trait is absent.

Proximal: The crown is shorter than the first molar and the palatal root has less diverence. The roots tend to remain within the crown profile.

Occlusal: The distolingual cusp is smaller on the second than on the first molar. When it is much reduced in size, the crown outline is described as 'heart-shaped.' The Carabelli trait is usually absent. The order of cusp size, largest to smallest, is the same as the first but is more exaggerated: mesiolingual, mesiobuccal, distobuccal, and distolingual.

Contact Points; Height of Curvature: Both mesial and distal contacts tend to be centered buccolingually below the marginal ridges. Since themolars become shorter, moving from first to this molar, the contacts tend to appear more toward the center of the proximal surfaces.

Roots: There are three roots, two buccal and one lingual. The roots are less divergent than the first with their apices usually falling within the crown profile. The buccal roots tend to incline to the distal.

Note: The distolingual cusp is the most variable feature of this tooth. When it is large, the occlusal is somewhat rhomboidal; when reduced in size the crown is described as triangual or 'heart-shaped.' At times, the root may be fused.

FORMATION OF THE ROOT AND ITS ROLE IN ERUPTION

- As dentin and enamel is deposited the shape of the future crown appears.

- The cells just superficial to the horizontal diaphragm start to proliferate and grow pushing the horizontal diaphragm down into the mesenchym.

- This forms a tube.

- This tube is the epithelial root sheath of Hertwig's.

- The mesenchym cells lying inside the tube nearest to the epithelial root sheath are induced to differentiate into odontoblasts, which then start to deposit dentin.

- After the first dentin of the root has been laid down the inner epithelial cells of the sheath start to deposit an enameloid substance called intermediate cementum.

- The root sheath cells then separate from the intermediate cementum and breaks up in a network of epithelial strands.

- The mesenchym on the outside comes into contact with the intermediate cementum and differentiate into

cementoblasts, which will deposit the cementum.

- This cementum traps the collagenic fibres, of the periodontal ligament, which are also formed.

- Epithelium of the root sheath persists as epithelial rests of Malassez. Because the epithelium of the root sheath forms from enamel epithelium it can develop into ameloblasts which will deposit enamel pearls.

- There is little space for the root to develop.

- To create space the crown is pushed out.

1. Errors in development. These are usually genetic.

a. Variability of the individual teeth. In general, the teeth most distal in any class are the most variable.

b. Partial or total anodontia. missing teeth in children,

c. Supernumerary teeth.

d. Microdontia

e. Macrodontia

F. Microdontia

2. Errors in skeletal alignment. Malpositioned jaws disrupt normal tooth relationships.

3. Soft tissue problems.

-Ocasionally, the proper eruption of a tooth is prevented by fibrous connective tissue over the crown of the tooth.

-In the mixed dentition, the deciduous second molars have a special importance for the integrity of the permanent dentition. Consider this: The first permanent molars at age six years erupt distal to the second deciduous molars.

-Permanent posterior teeth exhibit physiological mesial drift, the tendency to drift mesially when space is available. If the deciduous second molars are lost prematurely, the first permanent molars drift anteriorly and block out the second premolars.

An incisor diastema may be present. The plural for diastema is diastemata.

-Important: The deciduous anteriors--incisors and canines are narrower than their permanent successors mesiodistally.

-Important: The deciduous molars are wider that their permanent successors mesiodistally.

-This size difference has clinical significance. The difference is called the leeway space.

The leeway space in the lower arch is approximately 3.4 mm.

-The leeway space in the upper arch is approximately 1.8 mm. In normal development, the leeway space is taken up by the mesial migration of the first permanent molars.