FORMATION OF THE PERMANENT DENTITION

Twenty deciduous tooth buds are formed initially.
Proliferative activity of the dental lamina during the bell stage that leads to formation of permanent tooth buds (cap stage) lingual of each deciduous tooth germ.
Molars have no predecessors; they are formed by posterior proliferation of the dental lamina.

HARD TISSUE FORMATION

Hard tissue formation starts at the late stages of the bell stage.
Differentiatioin of cells into odontoblasts and ameloblasts.
The cells of the inner dental epithelium will become ameloblasts.
The cells of the dental papilla opposite to the inner dental epithelium will become odontoblasts.
Dentin is formed before enamel.
Dentin initiates the formation of enamel.

ROOT FORMATION

The root of the tooth is composed by dentin and cementum.
Dentinogenesis is initiated by the odontoblasts.
Odontoblasts are formed as epithelial cells continue to proliferate from the cervical loop as a double layer of cells known as Hertwig's root sheath.

TOOTH SHAPE

The shape of the crowns results from the interaction of inner dental epithelium and the dental papilla.
The cells of the inner dental epithelium have a programmed proliferation.
This internal program determines the tooth form.

The fate of the dental lamina

Rests of Serres
The rest of Serres are rests of the dental lamina identified in the gingival soft tissues.
They are round to ovoid aggregates of epithelial cells that have clear cytoplasm (glucogen rich).
They result from early breakup of the dental lamina during bell stage.

Rests of Malassez
The rests of Malassez result from breakup of the Hertwig's root sheath during root formation.
They can be identified in the periodontal ligament and are responsible for the development of radicular cysts.

INNERVATION OF THE DENTIN-PULP COMPLEX

1. Dentine Pulp
2. Dentin
3. Nerve Fibre Bundle
4. Nerve fibres

The nerve bundles entering the tooth pulp consist principally of sensory afferent fibers from the trigeminal nerve and sympathetic branches from the superior cervical ganglion. There are non-myelinated (C fibers) and myelinated (less than non, A-delta, A-beta) fibers. Some nerve endings terminate on or in association with the odontoblasts and others in the predentinal tubules of the crown. Few fibers are found among odontoblasts of the root.
In the cell-free zone one can find the plexus of Raschkow.

Age changes in the dentition

I. After the teeth have reached full occlusion, microscopic tooth movements occur to compensate for wear at the contact area (Mesial Drift) and occlusal surfaces (by Deposition of cementum at the root apex)

2. Attrition of incisal ridges and cusp tips may be so severe that dentin may become exposed and intrinsically stained

3. Secondary dentin may be formed in response to dental caries, trauma, and aging and result in decreased pulp size and tooth sensation

MANDIBULAR LATERAL INCISORS

The mandibular incisor is a little wider mesiodistal than the mandibular central incisor, and the crown is slightly longer from the incisal edge to the cervical line.

Facial Surface:-The facial surface is less symmetrical than the facial surface of the mandibular central incisor. The incisal edge slopes upward toward the mesioincisal angle, which is slightly less than 90°. The distoincisal angle is rounded. The mesial border is more nearly straight than the distal border.

Lingual Surface:- The incisal portion of the lingual surface is concave. The cingulum is quite large but blends in smoothly with the rest of the surface.

Root Surface:-The root is single and extremely flattened on its mesial and distal surfaces.

As root and cementum formation begin, bone is created in the adjacent area. Throughout the body, cells that form bone are called osteoblasts. In the case of alveolar bone, these osteoblast cells form from the dental follicle. Similar to the formation of primary cementum, collagen fibers are created on the surface nearest the tooth, and they remain there until attaching to periodontal ligaments.

Like any other bone in the human body, alveolar bone is modified throughout life. Osteoblasts create bone and osteoclasts destroy it, especially if force is placed on a tooth. As is the case when movement of teeth is attempted through orthodontics, an area of bone under compressive force from a tooth moving toward it has a high osteoclast level, resulting in bone resorption. An area of bone receiving tension from periodontal ligaments attached to a tooth moving away from it has a high number of osteoblasts, resulting in bone formation.

MANDIBULAR FIRST MOLAR

It is the first permanent tooth to erupt.

Facial Surface:- The lower first permanent molar has the widest mesiodistal diameter of all of the molar teeth. Three cusps cusps separated by developmental grooves make on the occlusal outline The mesiobuccal cusp is usually the widest of the cusps. The mesiobuccal cusp is generally considered the largest of the five cusps. The distal root is usually less curved than the mesial root.

Lingual: Three cusps make up the occlusal profile in this view: the mesiolingual, the distolingual, and the distal cusp which is somewhat lower in profile. The mesiobuccal cusp is usually the widest and highest of the three. A short lingual developmental groove separates the two lingual cusps

Proximal: The distinctive height of curvature seen in the cervical third of the buccal surface is called the cervical ridge. The mesial surface may be flat or concave in its cervical third . It is highly convex in its middle and occlusal thirds. The occlusal profile is marked by the mesiobuccal cusp, mesiolingual cusp, and the mesial marginal ridge that connects them. The mesial root is the broadest buccolingually of any of the lower molar roots. The distal surface of the crown is narrower buccolingually than the mesial surface. Three cusps are seen from the distal aspect: the distobuccal cusp, the distal cusp, and the distolingual cusp.

Occlusal There are five cusps. Of them, the mesiobuccal cusp is the largest, the distal cusp is the smallest. The two buccal grooves and the single lingual groove form the "Y" patern distinctive for this tooth

Roots :-The tooth has two roots, a mesial and a distal.

Contact Points; The mesial contact is centered buccolingually just below the marginal ridge. The distal contact is centered over the distal root, but is buccal to the center point of the distal marginal ridge.

Roots: Lower molars have mesial and distal roots. In the first, molar, the mesial root is the largest. It has a distal curvature. The distal root has little curvature and projects distally.