Embryonic development

The parotid derives from ectoderm
The sublingual-submandibular glands thought to derive from endoderm
Differentiation of the ectomesenchyme
Development of fibrous capsule
Formation of septa that divide the gland into lobes and lobules
The parotid develops around 4-6 weeks of embryonic lofe
The submandibular gland develops around the 6th week
The sublingual and the minor glands develop around the 8-12 week

MAXILLARY CUSPIDS (CANINE)

The maxillary cuspid is usually the longest tooth in either jaw. canines are considered the corner stones of the dental arch They are the only teeth in the dentition with a single cusp.

Facial Surface:- The facial surface of the crown differs considerably from that of the maxillary central or lateral incisors. In that the incisal edges of the central and lateral incisor are nearly straight, the cuspid has a definite point, or cusp.  There are two cutting edges, the mesioincisal and the distoincisal. The distoincisal cutting edge is the longer of the two. The developmental grooves prominent on the facial surface  extending two-thirds of the distance from the tip of the cusp to the cervical line.  The distal cusp ridge is longer than the mesial cusp ridge

Lingual Surface:  Distinct mesial and distal marginal ridges, a well-devloped cingulum, and the cusp ridges form the boundries of the lingual surface. The prominent lingual ridge extends from the cusp tip to the cingulum, dividing the lingual surface into mesial and distal fossae.

Proximal: The mesial and distal aspects present a triangular outline. They resemble the incisors, but are more robust--especially in the cingulum region

Incisal: The asymmetry of this tooth is readily apparent from this aspect. It usually thicker labiolingually than it is mesiodistally. The tip of the cusp is displaced labially and mesial to the central long axis of this tooth.

Root Surface:-The root is single and is the longest root in the arch. It is usually twice the length of the crown.

MAXILLARY SECOND MOLAR

The second molars are often called 12-year molars because they erupt when a child is about 12 years

Facial: The crown is shorter occluso-cervically and narrower mesiodistally whe compared to the first molar. The distobuccal cusp is visibly smaller than the mesiobuccal cusp. The two buccal roots are more nearly parallel. The roots are more parallel; the apex of the mesial root is on line with the with the buccal developmental groove. Mesial and distal roots tend to be about the same length.

Lingual: The distolingual cusp is smaller than the mesiolingual cusp. The Carabelli trait is absent.

Proximal: The crown is shorter than the first molar and the palatal root has less diverence. The roots tend to remain within the crown profile.

Occlusal: The distolingual cusp is smaller on the second than on the first molar. When it is much reduced in size, the crown outline is described as 'heart-shaped.' The Carabelli trait is usually absent. The order of cusp size, largest to smallest, is the same as the first but is more exaggerated: mesiolingual, mesiobuccal, distobuccal, and distolingual.

Contact Points; Height of Curvature: Both mesial and distal contacts tend to be centered buccolingually below the marginal ridges. Since themolars become shorter, moving from first to this molar, the contacts tend to appear more toward the center of the proximal surfaces.

Roots: There are three roots, two buccal and one lingual. The roots are less divergent than the first with their apices usually falling within the crown profile. The buccal roots tend to incline to the distal.

Note: The distolingual cusp is the most variable feature of this tooth. When it is large, the occlusal is somewhat rhomboidal; when reduced in size the crown is described as triangual or 'heart-shaped.' At times, the root may be fused.

Transient structures during tooth development

Enamel knot: Thickening of the internal dental epithelium at the center of the dental organ.
Enamel cord: Epithelial proliferation that seems to divide the dental organ in two.
 

Review the role of these two structures
Enamel niche: It is an artifact that is produced during section of the tissue. It occurs because the dental organ is a sheet of proliferating cells rather than a single strand. It looks like a concavity that contains ectomesenchyme.

Enamel

Structural characteristics and microscopic features

a.  Enamel rods or prisms

(1) Basic structural unit of enamel.

(2) Consists of tightly packed hydroxyapatite crystals. Hydroxyapatite crystals in enamel are four times larger and more tightly packed than hydroxyapatite found in other calcified

tissues (i.e., it is harder than bone).

(3) Each rod extends the entire thickness of enamel and is perpendicular to the dentinoenamel junction (DEJ).
 

b. Aprismatic enamel

(1) The thin outer layer of enamel found on the surface of newly erupted teeth.

(2) Consists of enamel crystals that are aligned perpendicular to the surface.

(3) It is aprismatic (i.e., prismless) and is more mineralized than the enamel beneath it.

(4) It results from the absence of Tomes processes on the ameloblasts during the final stages of enamel deposition.

c. Lines of Retzius (enamel striae)

(1) Microscopic features

 (a) In longitudinal sections, they are observed as brown lines that extend from the DEJ to the

tooth surface.

 (b) In transverse sections, they appear as dark, concentric rings similar to growth rings in a tree.
 

(2) The lines appear weekly during the formation of enamel.
 

(3) Although the cause of striae formation is unknown, the lines may represent appositional or incremental growth of enamel. They may also result from metabolic disturbances of ameloblasts.

(4) Neonatal line

(a) An accentuated, dark line of Retzius that results from the effect of physiological changes

on ameloblasts at birth.

(b) Found in all primary teeth and some cusps of permanent first molars.

d. Perikymata

(1) Lines of Retzius terminate on the tooth surface in shallow grooves known a perikymata.

(2) These grooves are usually lost through wear but may be observed on the surfaces of developing teeth or nonmasticatory surfaces of formed teeth.
 

e. Hunter-Schreger bands

(1) Enamel rods run in different directions. In longitudinal sections, these changes in direction result in a banding pattern known as HunterSchreger bands.

(2) These bands represent an optical phenomenon of enamel and consist of a series of  alternating dark and light lines when the section is viewed with reflected or polarized

light.

f. Enamel tufts

(1) Consist of hypomineralized groups of enamel rods.

(2) They are observed as short, dark projections found near or at the DEJ.

(3) They have no known clinical significance.

g. Enamel lamellae
 

(1) Small, sheet-like cracks found on the surface of enamel that extend its entire thickness.

(2) Consist of hypocalcified enamel.

(3) The open crack may be filled with organic material from leftover enamel organ components, connective tissues of the developing tooth, or debris from the oral cavity.

(4) Both enamel tufts and lamellae may be likened to geological faults in mature enamel.
 

h. Enamel spindle
 

(1) Remnants of odontoblastic processes that become trapped after crossing the DEJ during the differentiation of ameloblasts.
 

(2) Spindles are more pronounced beneath the cusps or incisal edges of teeth (i.e., areas where occlusal stresses are the greatest).
 

PULP

Coronal

Occupies and resembles the crown,

Contains the pulp horns

It decreases in size with age

Radicular

Occupies roots

Contains the apical foramen

It decreases in size with age

Accessory apical canals

PULP FUNCTIONS

Inductive: The pulp anlage initiates tooth formation and probably induces the dental organ to become a particular type of tooth.

Formative: Pulp odontoblasts develop the organic matrix and function in its calcification.

Nutritive: Nourishment of dentin through the odontoblasts.

Protective: Sensory nerves in the tooth respond almost always with PAIN to all stimuli (heat, cold, pressure, operative procedures, chamical agents).

Defensive or reparative: It responds to irritation by producing reparative dentin. The response to stimuli is inflammation.

 Histologically the pulp consists of delicate collagen fibers, blood vessels, lymphatics, nerves and cells. A histologic section of the pulp reveals four cellular zones:

Odontoblastic

Cell-free (Weil)

Cell-rich

Pulp core